Department Head: 

Prof. Dr. Alexandra-Maria Klein
phone: +49 (0)761 203-67770


Frau Ilona Winkler
phone: +49 (0)761 203-3635
fax: +49 (0)761 203-3638


Chair of Nature Conservation
& Landscape Ecology
University of Freiburg
Tennenbacher Str. 4
D-79106 Freiburg

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Prof. Alexandra-Maria Klein

Department Head Chair of Nature Conservation and Landscape Ecology

Klein_2016Alexandra-Maria Klein
phone: +49 (0)761 203-67770
Room No.: 03043 office hours: Thursday 9:15 - 10:15




since 2013 Full professorship in Nature Conservation and Landscape Ecology, University of Freiburg, Germany
2010 - 2013 Full professorship in Ecosystem Functions at the Leuphana University of Lüneburg, Germany
2009 - 2010 Head of the Agroecology group in the Department of Crop Sciences, Georg-August University of Göttingen, Germany
2007 - 2008 Visiting Researcher at UC Berkeley, Davis, Stanford, USA
2005 - 2007 Lecturer and Research Assistant, Institute of Agroecology, Georg-August University of Göttingen, Germany
2003 - 2005 Postdoc, Institute of Agroecology, project Bioteam "Evaluation of biological diversity of land-use systems in a mega-diverse region of Ecuador" financed by the BMBF (Federal Ministry of Education and Research)
2000 - 2003 Ph.D in Agroecology and Zoology at Institute of Agroecology, Germany and IPB Bogor, Indonesia (certificate Dr. sc. agr.)
1998 - 1999 Diploma in Biology with specialisation in tropical agriculture and pollination biology at University of Göttingen, Germany (certificate: diploma in biology)
1997Professional courses, work experience: Practical course and work experience in tropical and subtropical agroecology at the ecological farm of José Lutzenberger in Brazil; Tropical study field course in Bolivia in botany and geography
1993 - 1999 Study of biology (final examines in botany, tropical and subtropical plant science, conservation science, microbiology) and working at the Institute of Tropical and Subtropical Sciences



selected publications


  • Staniczenko, P., Lewis, O. Tylianakis, J., Albrecht, M., Coudrain, V., Klein, A.M. & Reed-Tsochas, F. (2017): Predicting the effect of habitat modification on networks of interacting species. Nature Communications, (accepted).
  • Schuldt, A., Fornoff, F., Bruelheide, H., Klein, A.M. & Staab, M. (2017): Tree species richness attenuates the positive relationship between mutualistic ant-hemipteran interactions and leaf chewer herbivory. Proceedings of the Royal Society B: Biological Sciences, (accepted).
  • Pufal, G., Steffan-Dewenter, I. & Klein, A.M. (2017): Crop pollination services at the landscape scale. Current Opinion in Insect Science 21: 91-97. Link
  • Schuldt, A., Bruelheide, H., Assmann, T., Buscot, F., Erfmeier, A., Klein, A.M., Ma, K., Scholten, T., Staab, M., Wirth, C., Zhang, J. & Wubet, T. (2017): Belowground top-down and aboveground bottom-up effects structure multitrophic communities in a biodiverse forest. Scientific Reports 7: 4222. doi: 10.1038/s41598-017-04619-3. Link
  • Staab, M., Fornoff, F., Klein, A.M. & Blüthgen, N. (2017): Ants at plant wounds - A little-known trophic interaction with evolutionary implications for ant-plant interactions. American Naturalist. online first. doi: 10.1086/692735. Link
  • Lichtenberg, E.M., Kennedy, C.M., Kremen, C., Batary, P., Berendse, F., Bommarco, R., Bosque-Perez, N.A., Carvalheiro, L.G., Snyder, W.E., William, Williams, N.M., Winfree, R., Astrom, S., Benjamin, F., Brittain, C., Chaplin-Kramer, R., Clough, Y., Connelly, H., Danforth, B., Diekotter, T., Eigenbrode, S.D., Ekroos, J., Elle, E., Freitas, B.F., Fukuda, Y., Gaines-Day, H.R., Gratton, C., Holzschuh, A., Isaacs, R., Isaia, M., Jha, S., Jonason, D., Jones, V.P., Klatt, B., Klein, A.M., Krauss, J., Letourneau, D.K., Macfadyen, S., Mallinger, R.E., Martin, E.A., Martinez, E., Memmott, J., Morandin, L., Neame, L., Otieno, M.,. Park, M.G., Pfiffner, L., Pocock, M., Ponce, C., Potts, S.G., Poveda, K., Ramos, M., Rosenheim, J.A., Rundlof, M., Sardinas, H., Saunders, M.E., Schon, N.L., Sciligo, A.R., Sidhu, C.S., Steffan-Dewenter, I., Tscharntke, T., Vesely, M., Weisser, W.W., Wilson, J.K. & Crowder, D.W. (2017): A global synthesis of the effects of diversified farming systems on arthropod diversity within fields and across agricultural landscapes. Global Change Biology. online first. doi: 10.1111/gcb.13714. Link
  • Ma, K., He, J.S., Bruelheide, H., Klein, A.M., Liu, X. & Schmid, B. (2017): Biodiversity–ecosystem functioning research in Chinese subtropical forests. Journal of Plant Ecology 10: 1-3. Link
  • Staab, M., Methorst, J., Peters, J., Blüthgen, N. & Klein, A.M. (2017): Tree diversity and nectar composition affect arthropod visitors on extrafloral nectaries in a diversity experiment. Journal of Plant Ecology 10: 201-212 . Link
  • Fornoff, F., Klein, A.M., Hartig, F., Benadi, G., Venjakob, C., Schaefer, M. & Ebeling, A. (2017): Functional flower traits and their diversity drive pollinator visitation. Oikos 126: 1020-1030. Link
  • Becker, N., Schön, S., Klein, A.M., Ferstl, I., Kizgin, A., Tannich, E., Kuhn, C., Pluskota, B. & Jöst, A. (2017): First mass development of Aedes albopictus (Diptera: Culicidae)—its surveillance and control in Germany. Parasitology Research 116: 847-858. Link
  • Drescher, N., Klein, A.M., Neumann, P., Yañez, O. & Leonhardt, S. (2017): Inside honeybee hives: Impact of natural propolis on the ectoparasitic mite Varroa destructor and viruses. Insects 8: 15. doi:10.3390/insects8010015. Link



  • Eardley, C., Freitas, B. M., Kevan, P.G., Rader, R., Gikungu, M., Klein, A.M., Maus, C., Meléndez-Ramírez, V. Palni, L.M.S., Vergara, C.H., & Wiantoro S. (2016): Chapter 1: Background to pollinators, pollination and food production. In: Potts, S.G., Imperatriz-Fonseca, V.L., & Ngo, H.T. (eds) Assessment report of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services on pollinators, pollination and food production. Secretariat of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services, Bonn, Germany. Link
  • Meyer, S.T., Ebeling, A., [..], Hudewenz, A., Klein, A.M, et al. & Weisser, W.W. (2016): Effects of biodiversity strengthen over time as ecosystem functioning declines at low and increases at high biodiversity. Ecosphere 7: e01619. Link
  • Gossner, M.M., Lewinsohn, T.M., Kahl, T., Grassein, F., Boch, S., Prati, D., Birkhofer, K., Renner, S.C., Sikorski, J., Wubet, T., Arndt, H., Baumgartner, V., Blaser, S., Blüthgen, N., Börschig, Buscot, F., C., Diekötter, T., Jorge, L.R., Jung, K., Keyel, A.C., Klein, A.M., Klemmer, S., Krauss, J., Lange, M., Müller, J., Overmann, J., Pašalić, E., Penone, C., Perović, D.J., Purschke, O., Schall, P., Socher, S.A., Sonnemann, I., Tschapka, M., Tscharnke, T., Türke, M., Venter, P.C., Weiner, C.N., Werner, M., Wolters, V., Wurst, S., Westphal, C., Fischer, M., Weisser, W.W. & Allan, E. (2016): Land-use intensification causes multitrophic homogenization of grassland communities. Nature 540: 266-269. Link
  • Maneke, A. & Klein, A.M. (2016): Untersuchungen zur Konkurrenz zwischen Honigbienen und Wildbienen in der Lüneburger Heide. Naturwissenschaftlicher Verein für das Fürstentum Lüneburg von 1851 e.V. 46: 121-132. Link
  • Boch, S., Allan, E., Birkhofer, K., Bossdorf, O., Blüthgen, N., Christ-Breulmann, S., Diekötter, T., Dormann, C.F., Gossner, M.M., Hallmann, C., Hemp, A., Hölzel, N., Jung, K., Klaus, V.H., Klein, A.M., Kleinbecker, T., Lange, M., Müller, J., Hacke, H., Prati, D., Renner, S.C., Rothenwörer, C., Schall, P., Schluze, E.D., Socher, S.A., Tscharntke, T., Türke, M., Weiner, C.N., Weisser, W.W., Westphal, C., Wolters, V., Wubet, T. & Tischer, M. (2016): Extensive und jährlich wechselnde Nutzungsintensität fördert den Artenreichtum im Grünland. ANLiegen Natur 38: 86-95. Link
  • Walmsley, D., Bailis, R. & Klein, A.M. (2016): A global synthesis of Jatropha cultivation: Insights into land use change and management practices. Environmental Science & Technology 50: 8993-9002. Link
  • Staab, M., Bruelheide, H., Durka, W., Michalski, S., Purschke, O., Zhu C.D. & Klein, A.M. (2016): Tree phylogenetic diversity promotes host-parasitoid interactions. Proceedings of the Royal Society B: Biological Sciences 283: 20160275. Link
  • Plath, M., Moser, C., Bailis, R., Brandt, P., Hirsch, H., Klein, A.M., Walmsley, D. & von Wehrden, H. (2016): A novel bioenergy feedstock in Latin America? Cultivation potential of Acrocomia aculeata under current and futur conditions. Biomass and Bioenergy 91: 186-195. Link
  • Klein, A.M. (2016): Fliegen, Motten, Wanzen. Honig- und Wildbienen haben viele Helfer, doch ersetzbar sind sie nicht. ADIZ - Die Biene – Imkerfreund April 2016, pp. 34-36. Link
  • Motzke, I., Klein, A.M., Saleh, S., Wanger, T.C. & Tscharntke, T. (2016): Habitat management on multiple spatial scales can enhance bee pollination and crop yield in tropical homegardens. Agriculture, Ecosystems & Environment 223: 144-151. Link
  • Dudenhöffer, J.H., Pufal, G., Roscher, C., & Klein, A.M. (2016): Plant density can increase invertebrate post-dispersal seed predation in an experimental grassland community. Ecology and Evolution 6: 3796-3807. Link
  • Venjakob, C., Klein, A.M., Ebeling, A., Tscharntke, T. & Scherber, C. (2016): Plant diversity increases spatio-temporal niche complementarity in plant-pollinator interactions. Ecology and Evolution 6: 2249-2261. Link
  • Kaluza, B.F., Wallace, H., Heard, T.A., Klein, A.M. & Leonhardt, S.D. (2016): Urban gardens promote bee foraging over natural habitats and plantations. Ecology and Evolution 6: 1304-1316. Link
  • Soliveres, S., Manning, P., Prati, D., Gossner, M.M., Alt, F., Arndt, H., Baumgartner, V., Binkenstein, J., Birkhofer, K., Blaser, S., Blüthgen, N., Boch, S., Böhm, S., Börschig, C., Buscot, F., Diekötter, T., Heinze, J., Hölzel, N., Jung, K., Klaus, V.H., Klein, A.M., Kleinebecker, T., Klemmer, S., Krauss, J., Lange, M., Morris, E.K., Müller, J., Oelmann, Y., Overmann, J., Pašalić, E., Renner, S.C., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schöning, I., Schrumpf, M., Sikorski, J., Socher, S.A., Solly, E.F., Sonnemann, I., Sorkau, E., Steckel, J., Steffen-Dewenter, I., Stempfhuber, B., Tschapka, M., Türke, M., Venter, P., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Wolters, V., Wubet, T., Wurst, S., Fischer, M. & Allan, E. (2016): Locally rare species influence grassland ecosystem multifunctionality. Philosophical Transactions of the Royal Society B 371: . Link
  • Peixoto-Pereira, M.H., Pufal, G., Staab, M., Martins, C.F. & Klein, A.M. (2016): Diversity and specificity of host-natural enemy interactions in an urban-rural interface. Ecological Entomology 41: 241-252. Link
  • Garibaldi, L.A., Aizen, M.A., Cunningham, S.A., Harder, L.D. & Klein, A.M. (2016): Incremental contribution of pollination and other ecosystem services to agricultural productivity: effects of service quantity and quality. In: Gemmill-Herren, B. (ed.) Pollination services to Agriculture: Sustaining and enhancing a key ecosystem service (Issues in Agricultural Biodiversity). Routledge, Abingdon, pp. 33-42. Link
  • Rader, R., Bartomeus, I., Garibaldi, L., Garratt, M.P.D., Howlett, B., Winfree, R., Cunningham, S.A., Mayfield, M.M., Arthur, A.D., Andersson, G.K.S., Blanche, R., Bommarco, R., Brittain, C., Carvalheiro, L.G., Chacoff, N.P, Entling, M.H., Foully, B., Freitas, B.M., Gemmill-Herren, B., Ghazoul, J., Griffin, S., Gross, C.L., Herbertsson, L., Herzog, F., Hipólito, J., Jaggar, S., Jauker, F., Klein, A.M., Kleijn, D., Krishnan, S., Queiroz Lemos, C., Lindström, S.A.M., Mandelik, Y., Magalhães Monteiro, V., Nelson, W., Nilsson, L., Pattemore, D., de O. Pereira, N., Pisanty, G., Potts, S.G., Reemer, M., Rundlöf, M., Sheffield, C.S., Scheper, J., Schüepp, C., Smith, H.G., Stanley, D.A., Stout, K.C., Szentgyörgyi, H., Taki, H., Vergara, C.H., Viana, B.F. & Woyciechowski, M. (2016): Non-bee insects are important contributors to global crop pollination. Proceedings of the National Academy of Sciences 113:: 146-151. Link (highlighted in Nature: 10.1038/528011d) Link



  • Schuldt, A., Wubet, T., Buscot, F., Staab, M., Assmann, T., Böhnke-Kammerlander, M., Both, S., Erfmeier, A., Klein, A.M., Ma, K., Pietsch, K., Schultze, S., Wirth, C., Zhang, J., Zumstein, P. & Bruelheide, H. (2015): Multitrophic diversity in a biodiverse forest is highly nonlinear across spatial scales. Nature Communications 6: 10169. Link
  • Miczajka, V.L., Klein, A.M. & Pufal, G. (2015): Elementary school children contribute to environmental research as citizen scientists. PLOS ONE 10: e0143229. doi: 10.1371/journal.pone.0143229. Link
  • Klein, A.M. (2015): Was wäre wenn? Die Bedeutung von Bienen und anderen Bestäubern für unsere Ernährung. AZID - Die Biene - Imkerfreund, Dezember 2015, pp. 14-18. Link
  • Hudewenz, A. & Klein, A.M. (2015): Red mason bees cannot compete with honey bees for floral resources in a cage experiment. Ecology and Evolution 5: 5049-5056. Link
  • Gámez-Virués, S., Perovic, D., Gossner, M., Börschig, C., Blüthgen, N., de Jong, H., Simons, N., Klein, A.M., Krauss, J., Maier, G., Scherber, C., Steckel, J., Rothenwöhrer, C., Steffan-Dewenter, I., Weiner, C., Weisser, W.W., Werner, M., Tscharntke, T. & Westphal, C. (2015): Landscape simplification filters species traits and drives biotic homogenization. Nature Communications 6: 8568. Link
  • Pufal, G. & Klein, A.M. (2015): Spatial scale affects seed predation and dispersal in contrasting anthropogenic landscapes. Basic and Applied Ecology 16: 726-736. Link
  • Garibaldi, L., Bartomeus, I., Bommarco, R., Klein, A.M., Cunningham, S.A., Aizen, M., Boreux, V., Garratt, M., Carvalheiro, L.G., Kremen, C., Morales, C., Schüepp, C., Chacoff, N., Freitas, B., Gagic, V., Holzschuh, A., Klatt, B., Krewenka, K., Krishnan, S., Mayfield, M., Motzke, I., Otieno, M., Petersen, J., Potts, S.G., Ricketts, T.H., Rundlöf, M., Sciligo, A., Sinu, P.A., Steffan-Dewenter, I., Taki, H., Tscharntke, T., Vergara, C.H., Viana, B.F. & Woyciechowski, M. (2015): Trait matching of flower visitors and crops predicts fruit set better than trait diversity. Journal of Applied Ecology 52: 1436-1444. Link
  • Staab, M., Blüthgen, N. & Klein, A.M. (2015): Tree diversity alters the structure of a tri-trophic network in a biodiversity experiment. Oikos 124: 827-834. Link
  • Schäckermann, J., Pufal, G., Mandelik, Y. & Klein, A.M. (2015): Agro-ecosystem services and dis-services in almond orchards are differentially influenced by the surrounding landscape. Ecological Entomology 40: 12-21. Link
  • Baumgärtner, S., Klein, A.M., Thiel, D. & Winkler, K. (2015): Ramsey discounting of ecosystem services. Enviromental and Resource Economics 61: 273-296 Link
  • Kleijn, D., Winfree, R., Bartomeus, I., Carvalheiro, L.G., Henry, M., Isaacs, R., Klein, A.M., Kremen, C., M'Gonigle, L.K., Rader, R., Ricketts, T., Williams, N.M., Adamson, N.L., Ascher, J.S., Báldi, A., Batáry, P., Benjamin, F., Biesmeijer, J.C., Blitzer, E.J., Bommarco, R., Brand, M.R., Bretagnolle, V., Button, L., Cariveau, D.P., Chifflet, R., Colville, J.F., Danforth, B.N., Elle, E., Garratt, M.P.D., Herzog, F., Holzschuh, A., Howlett, B.G., Jauker, F., Jha, S., Knop, E., Krewenka, K.M., Le Féon, V., Mandelik, Y., May, E.A., Park, M.G., Pisanty, G., Reemer, M., Riedinger, V., Rollin, O., Rundlöf, M., Sardiñas, H.,S., Scheper, J., Sciligo, A.R., Smith, H.G, Steffan-Dewenter, I., Thorp, R., Tscharntke, T., Verhulst, J., Viana, B.F., Vaissière, B.E., Veldtman, R., Westphal, C. & Potts, S.G. (2015): Delivery of crop pollination services is an insufficient argument for wild pollinator conservation. Nature Communications 6: 7414. Link
  • Klein, A.M. (2015): Bienen und ihre Bedeutung für die Bestäubungsleistung von Nutzpflanzen. In: Lorenz, S., Stark, K. (eds.) Menschen und Bienen. Ein nachhaltiges Miteinander in Gefahr. Oekom-Verlag, München, pp. 27-36. Link
  • Motzke, I., Tscharntke, T., Wanger, T.C. & Klein, A.M. (2015): Pollination mitigates cucumber yield gaps more than pesticide and fertilizer use in tropical smallholder gardens. Journal of Applied Ecology 52: 261-269. Link
  • Hülsmann, M., von Wehrden, H., Klein, A.M. & Leonhardt, S.D. (2015):  Plant diversity and composition compensate for negative effects of urbanization on foraging bumble bees. Apidologie 46: 760-770. Link
  • Groeneveld, J. & Klein, A.M. (2015): Pennycress-corn double-cropping increases ground beetle diversity. Biomass and Bioenergy 77: 16-25. Link
  • Staab, M., Ohl, M., Zhu, C.D. & Klein, A.M. (2015): Observational natural history and morphological taxonomy are indispensable for future challenges in biodiversity and conservation. Communicative and Integrative Biology 8: e992745. doi: 10.4161/19420889.2014.992745. Link
  • Schäckermann, J., Mandelik, Y., Weiss, N., von Wehrden, N., & Klein, A.M. (2015): Natural habitat does not mediate vertebrate seed predation as an ecosystem dis-service to agriculture. Journal of Applied Ecology 52: 291-299. Link
  • Klein, A.M. (2015): Contribution of pollinator-mediated crops to nutrients in the human food supply. In: Potts, S., Biesmeijer, K., Bommarco, R., Breeze, T., Carvalheiro, L., Franzén, M., González-Varo, J.P., Holzschuh, A., Kleijn, D., Kunin, B., Lecocq, T., Lundin, O., Michez, D., Neumann, P., Nieto, A., Penev, L., Rasmont, P., Ratamäki, O., Riedinger, V., Roberts, S.P.M., Rundlöf, M., Scheper, J., Sørensen, P., Steffan-Dewenter, I., Stoev, P., Vilà, M. & Schweiger O., (2015): Status and trends of European pollinators. Key findings of the STEP project. Pensoft Publishers, Sofia, pp. 41-42. Link
  • Perović, D., Gámez-Virués, S., Börschig, C., Klein, A.M., Krauss, J., Steckel, J., Rothenwöhrer, C., Erasmi, S., Tscharntke, T. & Westphal, C. (2015): Configurational landscape heterogeneity shapes functional community composition of grassland butterfliesJournal of Applied Ecology 52: 505-513. Link
  • Manning, P., Gossner, M., Bossdorf, O., Allan, E., Prati, D., Blüthgen, N., Boch, S., Böhm, S.,  Börschig, C., Hölzel, N., Jung, K., Klaus, V., Klein, A.M., Kleinebecker, T., Krauss, J., Lange, M., Müller, J., Pašalić, E., Socher, S., Tschapka, M., Türke, M., Weiner, C., Werner, M., Gockel, S., Hemp, A., Renner, S., Wells, K., Buscot, F., Kalko, E., Linsenmair, K., Weisser, W.W. & Fischer, M. (2015): Grassland management intensification weakens the associations among the diversities of multiple plant and animal taxa. Ecology 6: 1492-1501. Link
  • Groeneveld, J., Lührs, H. & Klein, A.M. (2015): Pennycress double-cropping does not negatively impact spider diversity. Agriculture and Forest Entomology 17: 247-257. Link



  • Börschig, C., Klein, A.M. & Krauss, J. (2014): Effects of grassland management, endophytic fungi and predators on aphid abundance in two distinct regions. Journal of Plant Ecology 7: 490-498.  Link
  • Tallis, H. et al. (incl. Klein, A.M. & Pufal, G.) (2014): Working together: A call for inclusive conservation. Nature 515: 27-28. Link
  • Staab, M., Schuldt, A., Assmann, T., Bruelheide, H. & Klein, A.M. (2014): Ant community structure during forest succession in a subtropical forest in South-East China. Acta Oecologica 61: 32-40. Link
  • Assmann, T., Drees, C., Härdtle, W., Klein, A.M., Schuldt, A. & von Oheimb, G. (2014): Ökosystem und Biodiversität. In: Heinrichs, H., Michelsen, G. (eds.) Nachhaltigkeitswissenschaften, Springer Spektrum Berlin Heidelberg, Germany, pp. 147-175. Link
  • Klein, A.M., Boreux, V., Bauhus, J., Chappell, M.J., Fischer, J. & Philpott, S.M. (2014): Forest islands in an agricultural sea. In: Kettle, C.J. & Koh, L.P. (eds): Global Forest Fragmentation, CABI, Delémont, Schweiz, pp. 79-95. Link
  • Hudewenz, A., Pufal, G., Bögeholz, A.L. & Klein, A.M. (2014): Cross-pollination benefits differ among oilseed rape varieties. Journal of Agricultural Sciences 152: 770-778. Link
  • Chaplin-Kramer, R., Dombeck, E., Gerber, J., Knuth, K.A., Mueller, N.D., Mueller, M., Ziva,G. & Klein, A.M. (2014). Global malnutrition overlaps with pollinator-dependent micronutrient production. Proceedings of the Royal Society B: Biological Sciences 281: 20141799. Link, Science Interview
  • Pereira-Peixoto, H.M., Pufal, G., Martins, C.F. & Klein, A.M. (2014): Spillover of trap-nesting bees and wasps in an urban-rural interface. Journal of Insect Conservation 18: 815-825. Link
  • Garibaldi, L.A., Carvalheiro, L.G., Leonhardt, S.D., Aizen, M.A., Blaauw, B.R., Isaacs, R., Kuhlmann, M., Kleijn, D., Klein, A.M., Kremen, C., Morandin, L., Scheper, J. & Winfree, R. (2014): From research to action: practices to enhance crop yield through wild pollinators. Frontiers in Ecology and the Environment 12: 439-447. Link, suppl.
  • Wanger, T.C., Tscharntke, T., Schroth, G. & Klein, A.M. (2014): Cocoa shortfall: Pollination curbs climate risk to cocoa.  Nature 511: 155. Link
  • Schäckermann, J., Weiss, N., von Wehrden, H. & Klein, A.M. (2014): High trees increase sunflower seed predation by birds in an agricultural landscape of Israel. Frontiers in Ecology and Evolution 2: 35. doi: 10.3389/fevo.2014.00035. Link
  • Staab, M., Ohl, M., Zhu, C.D. & Klein, A.M. (2014): A unique nest-protection strategy in a new species of spider wasp. PLOS ONE 9: e101592. doi: 10.1371/journal.pone.0101592. Link (highlighted in Nature 10.07.2014) Link
  • Staab, M., Schuldt, A., Assmann, T. & Klein, A.M. (2014): Tree diversity promotes predator but not omnivore ants in a subtropical Chinese forest. Ecological Entomology 39: 637-647. Link
  • Abbas, M., Klein, A.M., Ebeling, A., Oelmann, Y., Ptacnik, R., Weisser, WW. & Hillebrand, H. (2014): Plant diversity effects on pollinating and herbivorous insects can be linked to plant stoichiometry. Basic and Applied Ecology 15: 169-178. Link
  • Abson, D.J., von Wehrden, H., Baumgärtner, S., Fischer, J., Hanspach, J., Härdtle, W., Heinrichs, H., Klein, A.M., Lang, D.J., Martens, P. & Walmsley, D. (2014): Ecosystem services as a boundary object for sustainability. Ecological Economics 103: 29-37. Link
  • Allan, E., Bossdorf, O., Dormann, C.F., Prati, D., Gossner, M.M., Tscharntke, T., Blüthgen, N., Barto, K., Bellach, M., Birkhofer, K., Boch, S., Böhm, S., Börschig, C., Chatzinotas, A., Christ, S., Daniel, R., Diekoetter, T., Fischer, C., Friedl, T., Glaser, K., Hallman, C., Hodaĉ, L., Hölzel, N., Jung, K., Klein, A.M., Klaus, V., Kleinebecker, T., Krauss, J., Lange, M., Müller, J., Nacke, H., Pašalić, E., Rillig, M., Rothenwöhrer, C., Schall, P., Scherber, C., Schulze, W., Socher, S., Steckel, J., Steffan-Dewenter, I., Türke, M., Weiner, C., Werner, M., Westphal, C., Wolters, V., Wubet, T., Gockel, S., Gorke, M., Hemp, A., Renner, S.C., Schöning, I., Pfeiffer, S., König-Ries, B., Buscot, F., Linsenmair, K.E., Schulze, E.D., Weisser, W.W. & Fischer, M. (2014): Interannual variation in land-use intensity enhances grassland multidiversity. Proceedings of the National Academy of Sciences 111: 308-313. Link
  • Brittain, C., Kremen, C., Garbers, A. & Klein, A.M. (2014): Pollination and plant resources change the nutritional quality of almonds for human health. PLOS ONE 9: e90082. doi: 10.1371/journal.pone.0090082. Link
  • Bruelheide, H., Nadrowski, K., Assmann, T., Bauhus, J., Both, S., Buscot, F., Chen, X.Y., Ding, B., Durka, W., Erfmeier, A., Gutknecht, J.L.M., Guo, D., Guo, L.D., Härdtle, W., He, J. S., Klein, A.M., Kühn, P., Liang, Y., Liu, X., Michalski, S., Niklaus, P.A., Pei, K., Scherer-Lorenzen, M., Scholten, T., Schuldt, A., Seidler, G., Trogisch, S., von Oheimb, G., Welk, E., Wirth, C., Wubet, T., Yang, X., Yu, M., Zhang, S., Zhou, H., Fischer, M., Ma, K. & Schmid, B. (2014): Designing forest biodiversity experiments: general considerations illustrated by a new large experiment in subtropical China. Methods in Ecology and Evolution 5: 74-89. Link
  • Groeneveld, J.H. & Klein, A.M. (2014): Pollination of two oil-producing plant species: Camelina (Camelina sativa L. Crantz) and pennycress (Thlaspi arvense L.) double cropping in Germany. Global Change Biology Bioenergy 6: 242-251. Link
  • Klein, A.M., Hendrix, S.D., Clough, Y., Scofield, A. & Kremen, C. (2014): Interacting effects of pollination, water and nutrients on fruit tree performance. Plant Biology 17: 201-208. Link
  • Pisanty, G., Klein, A.M. & Mandelik, Y. (2014): Do wild bees complement honey bee pollination of confection sunflowers in Israel? Apidologie 45: 235-247. Link
  • Wanger, T.C., Darras, K., Bumrungsri, S., Tscharntke, T. & Klein, A.M. (2014): Bat pest control contributes to food security in Thailand. Biological Conservation 171: 220-223. Link



  • Allan, E., Weisser, W.W., Fischer, M., Schulze, E.D., Weigelt, A., Roscher, C., Baade, J., Barnard, R.L., Bessler, H., Buchmann, N., Buscot, F., Ebeling, A., Eisenhauer, N., Engels, C., Fergus, A.J.F., Gleixner, G., Gubsch, M., Habekost, M., Halle, S., Klein, A.M., König, S., Kowalski, E., Kreutziger, Y., Kertscher, I., Kummer, V., Kuu, A., Lange, M., Lauterbach, D., le Roux, X., Marquard, E., Migunova, V.D., Milcu, A., Müller, R., Mwangi, P., Niklaus, P.A., Oelmann, Y., Petermann, J.S., Poly, F., Rottstock, T., Rosenkranz, S., Sabais, A., Scherber, C., Scherer-Lorenzen, M. Scheu, S., Schmitz, M., Schumacher, J., Soussana, J.F., Steinbeiss, S., Schwichtenberg, G., Temperton, V., Tscharntke, T., Voigt, W., Wilcke, W., Wirth, C. & Schmid, B. (2013): A comparison of the strength of biodiversity effects across multiple functions. Oecologia 173: 223-237. Link
  • Börschig, C., Klein, A.M., von Wehrden, H. & Krauss, J. (2013): Traits of butterfly communities change from specialist to generalist characteristics with increasing land-use intensity. Basic and Applied Ecology 14: 547-554. Link
  • Brittain, C., Kremen, C., Hendrix, S.D., Thorp, R., Williams, N. & Klein, A.M. (2013): Almond orchards with wild insects have higher fruit set. Sacramento Valley Regional Almond Newsletter, p. 4.
  • Brittain, C., Williams, N., Kremen, C. & Klein, A.M. (2013): Synergistic effects of non-Apis bees and honey bees for pollination services. Proceedings of the Royal Society B. 280: 20122767. online first. doi: 10.1098/rspb.2012.2767. Link
  • Fischer, J., Brittain, C. & Klein, A.M. (2013): Biodiversity-friendly farming. In: Levin, S.A. (eds.): Encyclopedia of Biodiversity, 2nd Edition, Elsevier, New York. Link
  • Garibaldi, L.A., Steffan-Dewenter, I., Winfree, R., Aizen, M.A., Bommarco, R., Cunningham, S.A., Kremen, C., Carvalheiro, L.G., Harder, L.D., Afik, O., Bartomeus, I., Benjamin, F., Boreux, V., Cariveau, D., Chacoff, N.P., Dudenhöffer, J.H., Freitas, B.M., Ghazoul, J., Greenleaf, S., Hipólito, J., Holzschuh, A., Howlett, B., Isaacs, R., Javorek, S.K., Kennedy, C.M., Krewenka, K., Krishnan, S., Mandelik, Y., Mayfield, M.M, Motzke, I., Munyuli, T., Nault, B.A., Otieno, M., Petersen, J., Pisanty, G., Potts, S.G., Rader, R., Ricketts, T.H., Rundlöf, M., Seymour, C.L., Schuepp, C., Szentgyörgyi, H., Taki, H., Tscharntke, T., Vergara, C.H., Viana, B.F., Wanger, T.C., Westphal, C., Williams, N. & Klein, A.M. (2013): Wild pollinators enhance fruit set of crops regardless of honey bee abundance. Science 339: 1608-1611. Link
  • Hudewenz, A. & Klein, A.M. (2013): Competition between honey bees and wild bees and the role of nesting resources in a nature reserve. Journal of Insect Conservation 17: 1275-1283. Link
  • Kennedy, C.M., Lonsdorf, E., Neel, M.C., Williams, N.M., Ricketts, T.H., Winfree, R., Bommarco, R., Brittain, C., Burley, A.L., Cariveau, D., Carvalheiro, L.G., Chacoff, N.P., Cunningham, S.A., Danforth, B.N., Dudenhöffer, J.H., Elle, E., Gaines, H.R., Gratton, C., Greenleaf, S.S., Holzschuh, A., Isaacs, R., Javorek, S.K., Jha, S., Klein, A.M., Krewenka, K., Mandelik, Y., Mayfield, M.M., Morandin, L., Neame, L.A., Otieno, M., Park, M., Potts, S.G., Rundlöf, M., Saez, A., Steffan-Dewenter, I., Taki, H., Tuell, J.K., Viana, B.F., Veldtman, R., Westphal, C. & Kremen, C. (2013): A global quantitative synthesis of local and landscape effects on native bee pollinators in agroecosystems. Ecology Letters 15: 584-599. doi: 10.1111/ele.12028. Link
  • Klein, A.M. (2013): Teamarbeit: Honigbienen können nur gemeinsam mit wildlebenden Insekten die Bestäubung von Kulturpflanzen sichern. Deutsches Bienenjournal 5: 12-14.
  • Larson, B.M.H., Kueffer, C. and the ZIF Working Group (Brock, A., Edwards, P., Gross, M., Hall, M., Higgs, E., Jentsch, A., Jeschke, J.M., Klein, A.M. & Wynne, B.) (2013): Managing invasive species amidst high uncertainty and novelty. Trends in Ecology and Evolution 28: 255-256. Link
  • Leonhardt, S.D., Gallai, N., Garibaldi, L.A., Kuhlmann, M. & Klein, A.M. (2013): Economic gain, stability of pollination and bee diversity decrease from southern to northern Europe. Basic and Applied Ecology 14: 461-471. Link
  • Motzke, I., Tscharntke, T., Navjot, S., Klein, A.M. & Wanger, T. (2013): Ant seed predation, pesticide applications and farmers' income from tropical multi-cropping gardens. Agricultural and Forest Entomology 15: 245-254. Link
  • Pufal, G. & Klein, A.-M. (2013): Post-dispersal seed predation of three grassland species in a plant diversity experiment. Journal of Plant Ecology 6: 468-479. Link



  • Blitzer, E.J., Dormann, C.F., Holzschuh, A., Klein, A.M., Rand, T.A. & Tscharntke, T. (2012): Spillover of functionally important organisms between managed and natural habitats. Agriculture, Ecosystems and Environment 146: 34-43. Link
  • Brittain, C., Kremen, C. & Klein, A.M. (2012): Biodiversity buffers pollination from changes in environmental conditions. Global Change Biology 19: 540-547. Link
  • Ebeling, A., Klein, A.M., Weisser, W.W. & Tscharntke, T. (2012): Multitrophic effects of experimental changes in plant diversity on cavity-nesting bees, wasps, and their parasitoids. Oecologia 169: 453-465. Link
  • Hudewenz, A., Klein, A.M., Scherber, C., Stanke, L., Tscharntke, T., Vogel, A., Weigelt, A., Weisser, W.W. & Ebeling, A. (2012): Herbivore and pollinator responses to grassland management intensity along experimental changes in plant species richness. Biological Conservation 150: 42-52. Link
  • Klein, A.M., Brittain, C., Hendrix, S.D., Thorp, R., Williams, N. & Kremen, C. (2012): Wild pollination services to California almond rely on semi-natural habitat. Journal of Applied Ecology 49: 723-732. Link
  • Lüdeke-Freund, F., Walmsley, D., Plath, M., Wreesmann, J. & Klein, A.M. (2012): Sustainable plant oil production for aviation fuels - Assessment challenges and consequences for new feedstock concepts. Sustainability Accounting, Management and Policy Journal 3: 186-217. Link
  • Schleuning, M., Fründ, J., Klein, A.M., Abrahamczyk, S., Alarcón, R., Albrecht, M., Andersson, G.K.S., Bazarian, S., Böhning-Gaese, K., Bommarco, R., Dalsgaard, B., Dehling, D.M., Gotlieb, A., Hagen, M., Hickler, T., Holzschuh, A., Kaiser-Bunbury, C.N., Kreft, H., Morris, R.J., Sandel, B., Sutherland, W.J., Svenning, J.C., Tscharntke, T., Watts, S., Weiner, C.N., Werner, M., Williams, N.M. Winqvist, C., Dormann, C.F. & Blüthgen, N. (2012). Specialization of mutualistic interaction networks decreases towards tropical latitudes. Current Biology 22: 1925-1931. Link
  • Smukler, S., Philpott, S., Jackson, L., Klein, A.M, DeClerck, F., Winowiecki, L. & Palm, C. (2012): Ecosystem services in agricultural landscapes. In: Clerck, F.A.J., Ingram, J.C., Rio, C.R. (eds.): Integrating Ecology into poverty reduction. Springer Verlag, Berlin, Germany. Link
  • Tscharntke, T., Tylianakis, J.M., Rand, T.A., Didham, R.K., Fahrig, L., Batáry, P., Bengtsson, J., Clough, Y., Crist, T.O., Dormann, C.F., Ewers, R.M., Fründ, J., Holt, R.D., Holzschuh, A., Klein, A.M., Kleijn, D., Kremen, C., Landis, D.A., Laurance, W., Lindenmayer, D., Scherber, C., Sodhi, N., Steffan-Dewenter, I., Thies, C., van der Putten, W.H. & Westphal, C. (2012): Landscape moderation of biodiversity patterns and processes - eight hypotheses. Biological Reviews 87: 661-685. Link
  • Vanbergen, A.J., Ambrose, N., Aston, D., Biesmeijer, C.J., Bourke, Breeze, A.T., Brotherton, P., Brown, M., Chandler, D., Clook, M., Connolly, C.N., Costigan, P., Coulson, M., Cresswell, J., Dean, R. Dicks, L., Felicioli, A., Fojt, O., Gallai, N., Genersch, E., Godfray, C., Grieg-Gran, M. Halstead, A., Harding, D., Harris, B., Hartfield, C., Heard, M.S., Herren, B., Howarth, J., Ings, T., Kleijn, D., Klein, A.M., Kunin, W.E., Lewis, G., MacEwen, A., Maus, C., McIntosh, L., Millar, N.S., Neumann, P., Ollerton, J., Olschewski, R., Osborne, J.L., Paxton, R.J., Pettis, J., Phillipson, B., Potts, S.G., Pywell, R., Rasmont, P., Roberts, S., Salles, J.M., Schweiger, O., Sima, P., Thompson, H., Titera, D., Vaissiere, B., Van der Sluijs, J., Webster, S., Wentworth, J. & Wright, G.A. (2012): Insect pollinators: linking research and policy. Church House Conference Centre, Dean's Yard, Westminster, London, 14 February 2012. Workshop Report, pp. 35. Link



  • Blüthgen, N. & Klein, A.M. (2011): Functional complementarity and specialisation: the role of biodiversity in plant-pollinator interactions. Basic and Applied Ecology 12: 282-291. Link
  • Ebeling, A., Klein, A.M., & Tscharntke, T. (2011): Plant-flower visitor interaction webs: Temporal stability and pollinator specialization increases along an experimental plant diversity gradient. Basic and Applied Ecology 12: 300-309. Link
  • Eilers, E.J., Kremen, C., Smith Greenleaf, S., Garber, A.K. & Klein, A.M. (2011): Contribution of pollinator-mediated crops to nutrients in the human food supply. PLoS ONE 6: e21363. doi: 10.1371/journal.pone.0021363. Link
  • Fischer, J., Batáry, P., Bawa, K.S., Brussaard, L., Chappell, M.J., Clough, Y., Daily, G.C., Dorrough, J., Hartel, T., Jackson, L.E., Klein, A.M., Kremen, C., Kuemmerle, T., Lindenmayer, D.B., Mooney, H.A., Perfecto, I., Philpott, S.M., Tscharntke, T., Vandermeer, J., Wanger, T.C. & von Wehrden, H. (2011): Conservation: Limits of land sparing. Science 334: 393-395. Link
  • Garibaldi, L.A., Aizen, M.A., Klein, A.M., Cunningham, S.A. & Harder, L.D. (2011): Global growth and stability of agricultural yield decrease with pollinator dependence. Proceedings of the National Academy of Science 108: 5909-5914. Link
  • Garibaldi, L.A., Steffan-Dewenter, I., Kremen, C., Morales, J.M., Bommarco, R., Cunningham, S.A., Carvalheiro, L.G., Chacoff, N.P., Dudenhöffer, J.H., Greenleaf, S.S., Holzschuh, A., Isaacs, R., Krewenka, C., Mandelik, Y., Mayfield, M.M., Morandin, L.A., Potts, S.G., Ricketts, T.H., Szentgyörgyi, H., Viana, B.F., Westphal, C., Winfree, R. & Klein, A.M. (2011): Stability of pollination services decreases with isolation from natural areas despite honey bee visits. Ecology Letters 14: 1062-1072. Link
  • Garibaldi, L.A., Muchhala, N., Motzke, I., Bravo-Monroy, L, Olschewski, R., Klein, A.M. (2011): Services from plant-pollinator interactions in the Neotropics. In: Rapidel, B., De Clerck, F., LeCoq, J.F., Beer, J. (eds): Ecosystem services from agriculture and agroforestry - Measurement and payment. Earthscan publishing.
  • Klein, A.M. (2011): Plant-pollinator interactions in changing environments. Basic and Applied Ecology 12: 279-281. Link
  • Olschewski, R. & Klein, A.M. (2011): Ecosystem services between sustainability and efficiency. Sustainability: Science, Practice, & Policy 7: 69-73. Link
  • Potts, S.G., Biesmeijer, J.C., Bommarco, R., Felicioli, A., Fischer, M., Jokinen, P., Kleijn, D., Klein, A.M., Kunin, W.E., Neumann, P., Penev, L.D., Petanidou, T., Rasmont, P., Roberts, S.P.M., Smith, H.G., Sørensen, P.B., Steffan-Dewenter, I., Vaissière, B.E., Vilà, M., Vujić, A., Woyciechowski, M., Zobel, M., Settele, J. & Schweiger, O. (2011): Developing European conservation and mitigation tools for pollination services: approaches of the STEP (Status and Trends of European Pollinators) project. Journal of Apicultural Research 50: 152-164. Link
  • Teodoro, A.V., Munoz A., Tscharntke,T., Klein, A.M. & Tylianakis, J.M. (2011): Early succession arthropod community changes on experimental passion fruit plant patches along a land-use gradient in Ecuador during the dry season. Agriculture, Ecosystems and Environment 140: 14-19. Link



  • Clough, Y., Abrahamczyk, S., Adams, M.O., Anshary, A., Ariyanti, N., Betz, L., Buchori, D., Cicuzza, D., Darras, K., Dwi Putra, D., Fiala, B., Gradstein, S.R., Kessler, M., Klein, A.M., Pitopang, R., Sahari, B., Scherber, C., Schulze, C.H., Shahabuddin, Sporn, S., Stenchly, K., Tjitrosoedirdjo, S.S., Wanger, T.C., Weist, M., Wielgoss, A. & Tscharntke, T. (2010): Biodiversity patterns and trophic interactions in human-dominated tropical landscapes in Sulawesi (Indonesia): plants, arthropods and vertebrates. In: Tscharntke, T., Leuschner, C., Veldkamp, E., Faust, H., Guhardja, E. & Bidin, A. (eds.): Tropical rainforests and agroforests under global change. Environmental Science and Engineering Series. Springer Verlag, Berlin, Germany, pp. 15-71. Link
  • Olschewski, R., Klein, A.M. & Tscharntke, T. (2010): Economic trade-offs between carbon sequestration, timber production, and crop pollination in tropical forested landscapes. Ecological Complexity 7: 314-319. Link
  • Scherber, C., Eisenhauer, N., Weisser, W.W., Schmid, B., Voigt, W., Schulze, E.D., Roscher, C., Weigelt, Al, Allan, E., Beßler, H., Bonkowski, M., buchmann, N., Buscot, F., Clement, L.W., Ebeling, A., Engels, C., Fischer, M.S., Halle, S., Kertscher, I., Klein, A.M., Koller, R., König, S., Kowalski, E., Kummer, V., Kuu, A., Lange, M., Lauterbach, D., Middelhoff, C., Migunova, V.D., Milcu, A., Müller, R., Partsch, S., Petermann, J.S., Renker, C., Rottstock, T., Sabais, A., Scheu, S., Schuchmacher, J., Temperton, V.M. & Tscharntke, T. (2010): Bottom-up effects of plant diversity on biotic interactions in a biodiversity experiment. Nature 468: 553-556. Link
  • Teodoro, A.V., Sousa-Souto, L., Klein, A.M. & Tscharntke, T. (2010): Seasonal contrasts in the response of coffee ants to agroforestry shade-tree management. Environmental Entomology 39: 1744-1750. Link
  • Tscharntke, T., Dormann, C., Holzschuh, A., Klein, A.M. & Thies, C. (2010): Bedeutung und Management der Bestäubung in Kulturlandschaften. In: Hotes, S. and Wolters, V. (eds.): Fokus Biodiversität - Wie Biodiversität in der Kulturlandschaft erhalten und nachhaltig genutzt werden kann. Oekom Verlag, München, Germany. Link
  • Veddeler, D., Tylianakis, J., Tscharntke, T. & Klein, A.M. (2010): Natural enemy diversity reduces temporal variability in wasp but not bee parasitism. Oecologia 162: 755-762. Link



  • Aizen, M.A., Garibaldi, L.A., Cunningham, S.A. & Klein, A.M. (2009): How much does agriculture depend on pollinators? Lessons from long-term trends in crop production. Annals of Botany 103: 1579-1588. Link
  • Eilers, E.J. & Klein, A.M. (2009): Landscape context and management effects on an important insect pest and its natural enemies in almond. Biological Control 51: 388-394. Link
  • Garibaldi, L.A., Aizen, A.A., Cunningham, S.A. & Klein, A.M. (2009): Pollination shortage and global crop production: looking at the whole spectrum of pollinator dependency. Communicative and Integrative Biology 2: 37-39. Link
  • Griffin, J.N., O'Gorman, E.J., Emmerson, M.C., Jenkins, S.R., Klein, A.M., Loreau & M., Symstad, A. (2009): Biodiversity and the stability of ecosystem functioning. In: Naeem S., Bunker, D.E., Hector, A., Loreau, M., Perrings, C. (eds.): Biodiversity, ecosystem functioning, and human wellbeing - an ecological and economic perspective. Oxford University Press, pp. 78-93. Link
  • Klein, A.M. (2009): Nearby rainforest promotes coffee pollination by increasing spatio-temporal stability in bee species richness. Forest Ecology and Management 258: 1838-1845. Link
  • Klein, A.M., Müller, C., Hoehn, P. & Kremen, C. (2009): Understanding the role of species richness for crop pollination services. In: Naeem, S., Bunker, D.E., Hector, A., Loreau, M., Perrings, C. (eds.): Biodiversity, ecosystem functioning, and human wellbeing - an ecological and economic perspective. Oxford University Press, pp. 195-208. Link
  • Teodoro, A.V., Klein, A.M. & Tscharntke T. (2009): Temporally mediated responses of the diversity of coffee mites to agroforestry management. Journal of Applied Entomology 133: 659-665. Link
  • Teodoro, A.V., Tscharntke, T. & Klein, A.M. (2009): From the lab to the field: contrasting effects of multitrophic interactions and agroforestry management on coffee pest densities. Entomologia Experimentalis et Applicata 131: 121-129. Link



  • Aizen, M.A., Garibaldi, L.A., Cunningham, S.A. & Klein, A.M. (2008): Long-term global trends in crop yield and production reveal no current pollination shortage but increasing pollinator dependency. Current Biology 18: 1572-1575. Link
  • Ebeling, A., Klein, A.M., Schumacher, J., Weisser, W. & Tscharntke, T. (2008): How does plant richness affect structure and stability of pollinator communities? Oikos 117: 1808-1815. Link
  • Klein, A.M., Cunningham, S.A., Bos, M. & Steffan-Dewenter, I. (2008): Advances in pollination ecology from tropical plantation crops. Ecology 89: 935-943. Link
  • Klein, A.M., Olschewski, R. & Kremen, C. (2008): The ecosystem service controversy: is there sufficient evidence for a 'pollination paradox'? GAIA 17: 12-16. Link
  • Kremen, C., Daily, G.C., Klein, A.M. & Scofield, D. (2008): Inadequate assesment of the ecosystem service rationale for conservation: a reply to Ghazoul. Conservation Biology 22: 795-798. Link
  • Lozada, T., de Koning, G.H.J, Kesseler, M., Klein, A.M. & Tscharntke, T. (2008): Geographical range size of tropical plants influence their response to anthropogenic activities. Diversity and Distributions 14: 59-68. Link
  • Ricketts, T.H., Regetz, J., Steffan-Dewenter, I., Cunningham, S.A., Kremen, C., Bogdanski, A., Gemmill-Herren, B., Greenleaf, S.S., Klein, A.M., Mayfield, M.M., Morandin, L.A., Ochieng, A. & Viana, B.F. (2008): Landscape effects on crop pollination services: are there general patterns? Ecology Letters 11: 499-515. Link
  • Teodoro, A., Klein, A.M. & Tscharntke, T. (2008): Environmentally mediated coffee pest densities in relation to agroforestry management, using hierarchical partitioning analyses. Agriculture, Ecosystems and Environment 125: 120-126. Link
  • Tylianakis, J.M, Rand, T.M., Kahmen, A., Klein, A.M., Buchmann, N., Perner, J. & Tscharntke, T. (2008): Resource heterogeneity moderates the biodiversity-function relationship in real world ecosystems. PloS Biology 6: 947-956. Link
  • Veddeler, D., Olschewski, R., Tscharntke, T. & Klein, A.M. (2008): The contribution of non-managed social bees to coffee production: new economic insights based on farm-scale yield data. Agroforestry Systems 73: 109-114. Link



  • Bos, M.M., Veddeler, D., Bogdansi, A., Klein, A.M., Tscharntke, T., Steffan-Dewenter, I. & Tylianakis, J. M. (2007): Caveates to quantifying ecosystem services. Fruit abortion blurs benefits from crop pollination. Ecological Applications 17: 1841-1849. Link
  • Klein, A.M., Vaissière, B.E., Cane, J., Steffan-Dewenter, I., Cunningham, S.A., Kremen, C. & Tscharntke, T. (2007): Importance of pollinators in changing landscapes for world crops. Proceedings of the Royal Society B 274: 303-313. Link
  • Kremen, C., Williams, N.M., Aizen, M.A., Gemmill-Herren, B., Le Buhn, G., Minckley, R., Packer, L., Potts, S.G., Roulston, T., Steffan-Dewenter, I., Vázquez, D.P., Winfree, R., Adams, L., Crone, E.E., Greenleaf, S.S., Keitt, T.H., Klein, A.M., Regetz, J. & Ricketts, T. (2007): Pollination and other ecosystem services produced by mobile organisms: a conceptual framework for the effects of land use change. Ecology Letters 10: 299-314. Link
  • Lozada, T., de Koning, G.H.J, Kesseler, M., Klein, A.M., Tscharntke, T. (2007): Geographical range size of tropical plants influence their response to anthropogenic activities. Diversity and Distributions 14: 59-68. Link
  • Lozada, T., de Koning, G.H.J., Marché, R., Klein, A.M. & Tscharntke, T. (2007): Tree recovery and seed dispersal by birds, comparing forest, agroforestry and abandoned agroforestry in coastal Ecuador. Perspectives in Plant Ecology, Evolution and Systematics 8: 115-128. Link
  • Olschewski, R., Tscharntke, T., Benítez, P.C., Schwarze, S., Klein, A.M. (2007): Economic evaluation of ecosystem services as a basis for stabilizing rainforest margins? The example of pollination services and pest management in coffee landscapes. In: Tscharntke, T., Leuschner, C., Zeller, M., Guhardja, E., Bidin, A. (eds.): The stability of tropical rainforest margins, linking ecological, economic and social constraints of land use and conservation. Springer Verlag, Berlin, pp. 265-278. Link
  • Priess, J. A., Mimler, M., Klein, A.M., Schwarze, S., Tscharntke, T. & Steffan-Dewenter, I. (2007): Linking deforestation scenarios to pollination services and economic returns in coffee agroforestry systems. Ecological Applications 17: 407-417.Link
  • Richter, A., Klein, A.M., Tscharntke, T. & Tylianakis, J. (2007): Abandonement of coffee agroforests increases insect abundance and diversity. Agroforestry Systems 69: 175-182. Link



  • Klein, A.M. (2006): A drop in pollinating species might endanger the production of many useful plants - La diminution du nombre d’insectes pollinisateurs met en péril la production de nom-breuses planted de culture - Rückgang von Bestäuberarten könnte die Produktion vieler Kulturpflanzen gefährden - Peligra la producción de muchas plantas de cultivo por el retroceso de especies polinizadora. Fruit World International 4: 90-94.
  • Klein, A.M. (2006): Was geht uns der Tod der Bienen an? Ist die Produktion und Vielfalt unserer Kulturpflanzen durch den Rückgang vieler Insekten gefährdet? Gastkommentar Artenvielfalt. Braunschweiger Zeitung vom 20.11.06, p. 4.
  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2006): Rainforest promotes trophic interactions and diversity of trap-nesting hymenoptera in tropical agroforestry. Journal of Animal Ecology 75: 315-323. Link
  • Olschewski, R., Tscharntke, T., Benitez, P.C., Schwarze, S. & Klein, A.M. (2006): Economic evaluation of pollination services comparing coffee landscapes in Ecuador and Indonesia. Ecology & Society 11. Link
  • Steffan-Dewenter, I. & Klein, A.M. (2006): Der ökologische Nutzen der Imkerei. Deutsches Bienenjournal 5: 4-5.
  • Steffan-Dewenter, I., Klein, A.M., Gaebele, V., Alfert, T. & Tscharntke, T. (2006): Bee diversity and plant-pollinator interactions in fragmented landscapes. In: Waser, N. & Ollerton, J. (eds.): Plant-pollinator interactions from specialization to generalization, pp. 387- 407. Link
  • Tylianakis, J. M., Klein, A.M., Lozada, T. & Tscharntke, T. (2006): Spatial scale of observation affects alpha, beta and gamma diversity of cavity-nesting bees and wasps across a tropical land-use gradient. Journal of Biogeography 33: 1295-1304. Link
  • Tylianakis, J. M., Tscharntke, T. & Klein, A.M. (2006): Diversity, ecosystem function and stability of parasitoid-host interactions across a tropical gradient of habitat modification. Ecology 87: 3047-3057. Link
  • Veddeler, D., Klein, A.M. & Tscharntke, T. (2006): Contrasting responses of bee communities to coffee flowering at different spatial scales. Oikos 112: 594-601. Link



  • Tscharntke, T., Klein, A.M., Kruess, A. & Steffan-Dewenter, I. (2005): Landscape perspectives on agricultural intensification and biodiversity - ecosystem service management. Ecology Letters 8: 857-874. Link
  • Tylianakis, J.M., Klein, A.M. & Tscharntke, T. (2005): Spatiotemporal variation in the diversity of hymenoptera across a tropical habitat gradient. Ecology 86: 3296-3302. Link



  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2004): Foraging trip duration and reproductive success of megachilid bees, eumenid wasps, and pompilid wasps in tropical agroforestry systems. Journal of Animal Ecology 73: 517-525. Link
  • Krauss, J., Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2004): Effects of habitat area, isolation, and landscape diversity on plant species richness of calcareous grasslands. Biodiversity and Conservation 13: 1427-1439. Link
  • Tylianakis, J.M., Veddeler, D., Lozada, T., López, R.M., Benítez, P., Klein, A.M., de Koning, G.H.J., Olschewski, R., Veldkamp, E., Navarrete, H., Onore, G. & Tscharntke, T. (2004): Biodiversity of land-use systems in coastal Ecuador and bioindication using trap-nesting bees, wasps, and their natural enemies. Lyonia 6: 8-15. Link



  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2003): Pollinator diversity and fruit set of Coffea canephora in relation to local and regional agroforestry management. Journal of Applied Ecology 40: 837-845. Link
  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2003): Fruit set of highland coffee increases with the diversity of pollinating bees. Proceedings of the Royal Society B 270: 955-961. Link
  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2003): Bee pollination and fruit set of Coffea arabica and C. canephora (Rubiaceae). American Journal of Botany 90: 153-157. Link
  • Krauss, J., Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2003): Effects of habitat area, isolation, and landscape diversity on plant species richness of calcareous grasslands. Biodiversity and Conservation 13: 1427-1439. Link
  • Tscharntke, T., Klein, A.M., Kruess, A., Steffan-Dewenter, I. & Thies, C. (2003): Biodiversität und Pflanzen-Insekten-Interaktionen in Kulturlandschaften. In: Gradstein, S.R., Willmann, R., Zizka, G. (eds.): Biodiverstiätsforschung, Die Entschlüsselung der Artenvielfalt in Raum und Zeit. Kleine Senckenberg-Reihe 45: 171-183. Link



  • Klein, A.M. (2002): Viele Bienenarten erzeugen viel Kaffee. Deutsches Bienenjournal 10: 24-25.
  • Klein, A.M., Steffan-Dewenter, I., Buchori, D. & Tscharntke, T. (2002): Effects of land-use intensity in tropical agroforestry systems on flower-visiting and trap-nesting bees and wasps. Conservation Biology 16: 1003-1014. Link
  • Klein, A.M., Steffan-Dewenter, I. & Tscharntke, T. (2002): Predator-prey ratios on cocoa in a land-use gradient in Indonesia. Biodiversity and Conservation 11: 683-693. Link
  • Tscharntke, T., Bürger, C., Eber, S., Gabriel, D., Grabe, H., Grönmeier, M., Holzschuh, A., Jahn, S., Klein, A.M., Klipfel, S., Kluth, S., Krauss, J., Kruess, A., Lehmann, K., Poveda, K., Roschewith, I., Schiele, S., Schmidt, M., Schulze, C. H., Steffan-Dewenter, I., Sundmacher, F., Thies, C., Veddeler, D. & Westphal, C. (2002): Pflanze-Insekt-Interaktionen und Biodiversität in der Agrarlandschaft. Forschungsmagazin Georgia Augusta: Vielfalt des Lebens - Biodiversität 1: 41-46. Link



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