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Department Head: 

Prof. Dr. Alexandra-Maria Klein
phone:+49 (0)761 203-67770



Mrs. Ilona Winkler
phone:+49 (0)761 203-3635
fax:+49 (0)761 203-3638


Chair of Nature Conservation
& Landscape Ecology
University of Freiburg
Tennenbacher Str. 4
D-79106 Freiburg




Dr. Felix Fornoff

20230117-P1174102-2.jpgFelix Fornoff
Web of Science ResearcherID: AAA-3776-2019
phone: +49 (0)761 203 -67790
Room No. 502



I am interested in the research areas of Biodiversity, Ecosystem-functioning, Community Ecology and Conservation. My research is centered on terrestrial ecosystems, especially insect-insect and insect-plant interactions. In this respect, I study the dependency of diversity and functioning on bottom up processes and in relation to human activities such as agricultural or urban land use. I am fascinated by ecosystems ranging for tropical forests to temperate grasslands and value the advantages of different approaches including replicated experimental manipulations, observational studies and citizen science approaches.

Current projects:


Besides theory, fieldwork, statistics and evaluation
I am interested in photography.
Mainly natural motifs, especially wild bees.
A small selection of my photos can also be found at:
Have fun with the fascination of nature!


Summer Semester:

  • M.Sc. Modul (51180) Formenkenntnisse, Biodiversität und Funktionen

Winter Semester:

  • M.Sc. Modul (13011) Wildbienenbestimmung





since 2018  Postdoc at the Chair of Nature Conservation and Landscape Ecology, Faculty of Environment and Natural Resources, University of Freiburg
2013 - 2017PhD thesis: "Multitrophic interaction and functional diversity in diversity experiments" at the Chair of Nature Conservation and Landscape Ecology, Faculty of Environment and Natural Resources, University of Freiburg
2008 - 2011Master of Biological Science, Faculty of Biology, University of Konstanz, Konstanz, Germany. Thesis in Biology: "Chemical Ecology of Aquatic Plant-Herbivore Interactions"
2005 - 2008Bachelor of Biological Science, Faculty of Biology, University of Konstanz, Konstanz, Germany. Thesis: "Top Radiotracking Movement and Activity of Neotropical Katydids"
2012Research internship, STRI-Intern, Barro Colorado Island, Panama. Research: "Ecology, structure and long term dynamics of neotropical forests"
2012Research internship, Stream and Riparian Research Laboratory, University of British Columbia, Vancouver, Canada. Research: "Top down effects on food web, community composition, detritivory and primary production in a temperate rainforest stream"
2011 - 2012Postgraduate research assistant, Faculty of Biology/Limnology, University of Konstanz, Konstanz, Germany.  Research: "Tracking resource reallocation in an aquatic macrophyte as response to herbivory using stable isotopes"
2011 - 2012Postgraduate research assistant, Faculty of Biology/Botanical Garden, University of Konstanz, Konstanz, Germany. Research: "Reviewing the ecology and control measures of the invasive Indian Balsam (Impatiens glandulifera) in Europe and Germany"
2009 - 2010Erasmus-Scholarship for International Studies at Cardiff University, Wales
2008Research internship, Smithsonian Tropical Research Institute, Gamboa, Panama. Research: "Foraging behavior of neotropical bats"
2007Field internship, Hellenic Ornithological Society, Pylos, Greece.  Field work: Conservation of the Chamaelio africanus







  • Wang, M.Q., Guo, S.K., Guo, P.F., Yang, J.J., Chen, G.A., Chesters, D., Orr, M., Niu, Z.Q.,  Staab, M., Chen, J.T., Li, Y., Zhou, Q.S., Fornoff, F., Shi, X., Li, S., Martini, M., Klein, A.M., Schuldt, A., Liu, X., Ma, K., Bruelheide, H., Luo, A. & Zhu, C.D. (2024): Multi-dimensionality of tree communities structure host-parasitoid networks and their phylogenetic composition. bioRxiv 2024.06.13.598779. Link

  • Fornoff, F., Lanner, J., Orr, M., Xie, T., Guo, S., Guariento, E., Tuerlings, T., Smagghe, G., Parys, K., Ćetković, A., Bila Dubaić, J., Geslin, B., Scharnhorst, V.S., Pachinger, B., Klein, A.M. & Meimberg, H. (2024): Home-and-away comparisons of life history traits indicate enemy release and founder effects of the solitary bee, Megachile sculpturalis. Basic and Applied Ecology 76: 69-79. Link
  • Klaus, F., Ayasse, M., Classen, A., […], Fornoff, F., [...], Klein, A.M. et al. & Pistorius, J. (2024): Improving wild bee monitoring, sampling methods, and conservation. Basic and Applied Ecology 75: 2-11. Link
  • Mupepele, A.C., von Königslöw, V., Bleile, A.M., Fornoff, F., Fründ, J. & Klein, A.M. (2023): Plant-pollinator interactions in apple orchards from a production and conservation perspective. BioRxiv (preprint). online first. doi: 10.1101/2023.11.20.567831. Link
  • Lindermann, L., Grabener, S., Fornoff, F., Hopfenmüller, S., Schiele, S., Stahl, J. & Dieker, D. (2023): Wildbienen und Wespen in Nisthilfen bestimmen: Ein Bestimmungschlüssel für Deutschland; Ratgeber. Braunschweig: Thünen-Institut für Biodiversität. doi: 10.3220/MX1685523077000. Link
  • Li, Y., Schmid, B., Schuldt, A. Li, S., Wang, M.Q., Fornoff, F., Staab, M., Guo, P.F., Anttonen, P., Chester, D., Bruelheide, H., Zhu, C.D., Ma, K. & Liu, X. (2023): Multitrophic arthropod diversity mediates tree diversity effects on primary productivity. Nature Ecology & Evolution 7: 832-840. Link
  • Fornoff, F., Halla, W., Geiger, S.; Klein, A.M. & Sann, M. (2023): DNA barcoding resolves quantitative multi-trophic interaction networks and reveals pest species in trap nests. Insect Conservation and Diversity 16: 725-731. Link
  • Georgi, M., Gärtner, S.M., Förschler, M.I., Buse, J., Fornoff, F., Ssymank, A. & Klein, A.M. (2023): Mulching time of forest meadows influences insect diversity. Insect Conservation and Diversity 16: 368-378. Link
  • Dürrbaum, E., Fornoff*, F., Scherber, C., Vesterinen, E.J. & Eitzinger*, B. (2023): Metabarcoding of trap nests reveals differential impact of urbanization on cavity nesting bee and wasp communities. Molecular Ecology 23: 6449-6460. Link *These authors contributed equally to the study


  • Tang, T., Zhang, N., Bongers, F., Staab, M., Schuldt, A., Fornoff, F., Lin, H.; Cavender-Bares, J., Hipp, A., Li, S., Liang, Y., Han, B., Klein, A.M., Bruelheide, H., Durka, W., Schmid, B., Ma, K. & Liu, X. (2022): Tree species and genetic diversity increase productivity via functional diversity and trophic feedbacks. eLife 11: e78703. Link
  • Georgi, M., Fornoff, F., Gärtner, S.M., Neitzel, S., Geist, A. & Klein, A.M. (2022): Timing and mulching frequency affected the number of nests of cavity-nesting wasps that hunt for aphids in forest meadows. Journal of Insect Conservation 26: 973-981. Link
  • Von Königslöw, V., Fornoff, F. & Klein, A.M. (2022): Wild bee communities benefit from temporal complementarity of hedges and flower strips in apple orchards. Journal of Applied Ecology 59: 2814-2824. Link
  • Ollerton, J., Trunschke, J., Havens, K., Landaverde-González, P., Keller, A., Gilpin, A.M., Rodrigo Rech, A., Baronio, G.J., Phillips, B.J., Mackin, C., Stanley, D.A., Treanore, E., Baker, E., Rotheray, E.L., Erickson, E., Fornoff, F., Brearley, F.Q., Ballantyne, G., Iossa, G., Stone, G.N., Bartomeus, I., Stockan, J.A., Leguizamón, J., Prendergast, K., Rowley, L., Giovanetti, M., de Oliveira Bueno, R., Wesselingh, R.A., Mallinger, R., Edmondson, S., Howard, S.R., Leonhardt, S.D., Rojas-Nossa, S.V., Brett, M., Joaqui, T., Antoniazzi, R., Burton, V.J., Feng, H.H., Tian, Z.X., Xu, Q., Zhang, C., Shi, C.L., Huang, S.Q., Cole, L.J., Bendifallah, L., Ellis, E.E., Hegland, S.J., Straffon Díaz, S., Lander, T.A., Mayr, A.V., Dawson, R., Eeraerts, M., Armbruster, W.S., Walton, B., Adjlane, N., Falk, S., Mata, L., Goncalves Geiger, A., Carvell, C., Wallace, C., Ratto, F., Barberis, M., Kahane, F., Connop, S., Stip, A., Sigrist, M.R., Vereecken, N.J., Klein, A.M., Baldock, K., & Arnold, S.E.J. (2022): Pollinator-flower interactions in gardens during the COVID-19 pandemic lockdown of 2020. Journal of Pollination Ecology 31: 87-96. Link
  • Klein, A.M., von Königslöw, V., Mupepele, A., Fornoff, F. & Wintermantel, D. (2022): Blühstreifen, Hecken und Spontanvegetation: Wie können wir Wildbienen im Obstanbau gezielt fördern? In: Eben, A. & Herz, A. (eds.) Online - Fachgespräch zum Thema „Biodiversität im Obstbau“. Journal für Kulturpflanzen 74: 11-18. Link
  • Von Königslöw, V., Fornoff, F. & Klein, A.M. (2022): Pollinator enhancement in agriculture: Comparing sown flower strips, hedges and sown hedge herb layers in apple orchards. Biodiversity and Conservation 31: 433-451. Link


  • Knuff, A., Nitsch, H., Stackelberg, N., Fornoff, F. & Schramek, J. (2021): FAKT-Blühmischungen zur Förderung von Bestäubern. Landinfo 3: 28-31. Link
  • Fornoff, F., Staab, M., Zhu, C.D. & Klein, A.M. (2021): Multi-trophic communities re-establish with canopy cover and microclimate in a subtropical forest biodiversity experiment. Oecologia 196: 289-301. Link
  • Guo, P.F., Wand, M.Q., Orr, M., Li, Y., Chen, J.T., Zhou, Q.S., Staab, M., Fornoff, F., Chen, G.H., Zhang, N.L., Klein, A.M., Zhu, C.D. (2021): Tree diversity promotes predatory wasps and parasitoids but not pollinator bees in a subtropical experimental forest. Basic and Applied Ecology 53: 134-142. Link


  • Roquer-Beni, L., Rodrigo, A., Arnan, X., Klein, A.M., Fornoff, F., Boreux, V. & Bosch, J. (2020): A novel method to measure hairiness in bees and other insect pollinators. Ecology and Evolution 10: 2979-2990. Link
  • Klein, A.M., Freitas, B.M., Bomfim, I.G.A., Boreux, V., Fornoff, F. & Oliveira, M.O. (2020): A Polinização Agrícola por Insetos no Brasil - Um Guia para Fazendeiros, Agricultores, Extensionistas, Políticos e Conservacionistas. Albert-Ludwigs University Freiburg, Nature Conservation and Landscape Ecology. FreiDok plus. online. doi: 10.6094/UNIFR/151237. 149p. Link
  • Klein, A.M., Freitas, B.M., Bomfim, I.G.A., Boreux, V., Fornoff, F. & Oliveira, M.O. (2020): Insect Pollination of Crops in Brazil - A Guide for Farmers, Gardeners, Politicians and Conservationists. Albert-Ludwigs University Freiburg, Nature Conservation and Landscape Ecology. FreiDok plus. online. doi: 10.6094/UNIFR/151200. 149p. Link 


  • Klein, A.M. & Fornoff, F. (2019): Die Bestäubungsprofis. In: Deutsches Bienen Journal - Sonderheft: Wildbienen Spezial. pp. 30-31. Link
  • Nitsch, H., Fornoff, F., Bauer, C., Sauer, N., Schäfer, L., Wunderlich, L., Claßen, A., Pabst, H., Schramek, J. & Klein, A.M. (2019): Ad hoc-Studie zur faunistischen Bewertung von FAKT-Blühmischungen. Institut für ländliche Strukturforschung, Frankfurt. interner Link, download
  • Schuldt, A., Ebeling, A., Kunz, M., Staab, M., Guimarães-Steinicke, C., Bachmann, D., Buchmann, N., Durka, W., Fichtner, A., Fornoff, F., Härdtle, W., Hertzog, L., Klein, A.M., Roscher, C., Schaller, J., von Oheimb, G., Weigelt, A., Weisser, W., Wirth, C., Zhang, J., Bruelheide, H. & Eisenhauer, N. (2019): Multiple plant diversity components drive consumer communities across ecosystems. Nature Communications 10: 1460. Link
  • Fornoff, F., Klein, A.M., Blüthgen, N. & Staab, M. (2019): Tree diversity increases robustness of multi-trophic interactions. Proceedings of the Royal Society B: Biological Sciences 286: 20182399. Link
  • Pauli, T., Castillo-Cajas, R.F., Rosa, P., Kukowka, S., Berg, A., van den Berghe, E., Fornoff, F., Hopfenmüller, S., Niehuis, M., Peters, R.S., Staab, M., Strumia, F., Tischendorf, S., Schmitt, T. & Niehuis, O. (2019): Phylogenetic analysis of cuckoo wasps (Hymenoptera: Chrysididae) reveals a partially artificial classification at the genus level and a species-rich clade of bee parasitoids. Systematic Entomology 44: 322-355. Link


  • Gutiérrez-Chacón, C., Fornoff, F., Ospina-Torres, R. & Klein, A.M. (2018): Pollination of granadilla (Passiflora ligularis) benefits from large wild insects. Journal of Economic Entomology 111: 1526-1534. Link
  • Cao, H., Klein, A.M., Zhu, C.-D., Staab, M., Durka, W., Fischer, M. & Fornoff, F. (2018): Intra- and interspecific tree diversity promotes multitrophic plant–Hemiptera–ant interactions in a forest diversity experiment. Basic and Applied Ecology 29: 89-97. Link
  • Klein, A.M., Fornoff, F., Mupepele, A.C., Boreux, V. & Pufal, G. (2018): Relevance of wild and managed bees for human well-being. Current Opinion in Insect Science 26: 82-88. Link


  • Cao, H.-X., Salle, J.L., Fornoff, F., Guo, P.-F. & Zhu, C.-D. (2017): Notes on Kocourekia Bouček (Hymenoptera: Eulophidae: Tetrastichinae) with description of a new species from China. Zootaxa 4317: 391–400. Link
  • Schuldt, A., Fornoff, F., Bruelheide, H., Klein, A.M. & Staab, M. (2017): Tree species richness attenuates the positive relationship between mutualistic ant-hemipteran interactions and leaf chewer herbivory. Proceedings of the Royal Society B: Biological Sciences 284: 20171489. Link
  • Staab, M., Fornoff, F., Klein, A.M. & Blüthgen, N. (2017): Ants at plant wounds - A little-known trophic interaction with evolutionary implications for ant-plant interactions. American Naturalist 190: 442-450. Link
  • Fornoff, F., Klein, A.M., Hartig, F., Benadi, G., Venjakob, C., Schaefer, M. & Ebeling, A. (2017): Functional flower traits and their diversity drive pollinator visitation. Oikos 126: 1020-1030. Link

2012 - 2015

  • Rothhaupt, K.-O., Fornoff, F. & Yohannes, E. (2015): Induced responses to grazing by an insect herbivore (Acentria ephemerella) in an immature macrophyte (Myriophyllum spicatum): an isotopic study. Ecology and Evolution 5: 3657-3665. Link
  • Fornoff, F. & Gross, E.M. (2014): Induced defense mechanisms in an aquatic angiosperm to insect herbivory. Oecologia 175: 173-185. Link
  • Fornoff, F., Dechmann, D.K.N. & Wikelski, M. (2012): Observation of movement and activity via radio-telemetry reveals diurnal behaviour of the neotropical katydid Philophyllia ingens (Orthoptera: Tettigonidae). Ecotropica 18: 27-34.

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